| A new species of the genus Liuixalus (Anura: Rhacophoridae) from southern China | |
| Yang JH1,2; Rao DQ3; Wang YY*1; wangyy@mail.sysu.edu.cn | |
| 2015 | |
| 发表期刊 | ZOOTAXA
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| 卷号 | 3990期号:2页码:247-258 |
| 合作性质 | 其它 |
| 摘要 | A new tree frog species of the genus Liuixalus was described from Heishiding Nature Reserve, Guangdong Province, China based on a combination of morphological characters and molecular phylogenetic analyses. Liuixalus feii sp. nov. is distinguished from its congeners by a combination of following characters: small size (SVL 16.2-17.6 mm in adult males and 18.0-18.7 mm in adult females); snout obtusely pointed; tympanum distinct, about half size of eye diameter; nostril closer to eye than to the tip of snout; fingers free of webbing; toe III longer than toe V; toes weakly webbed; tibiotarsal articulation reaching the naris or loreal; dorsal skin smooth and scattered with fine granulars; a subtle longitudinal median ridge present on dorsum; weak skin folds present on dorsal surface of body and thighs; supratympanic fold distinct and curved; ventral surface dull white with more or less irregular dark spots; iris bicolored. The new species appears to be forest-dependent and to date has only been found on the forest floor in primary forests at elevations between 350-800 m. Based on our molecular analyses, we consider Liuixalus catbaensis as a junior synonym of L. calcarius. Thus, with the description of the new species, the genus Liuixalus hitherto contains five recognized species, four of which are endemic to China. |
| 关键词 | Rhacophoridae Liuixalus New Species Taxonomy China |
| 资助者 | This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). |
| 收录类别 | 其他 |
| 语种 | 英语 |
| 资助者 | This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). ; This work was partially supported by the Natural Science Foundation of China (NSFC: Sun Yat-sen University biology field practice ability improve project base, No.J1210074) and the Project of Ministry of education of China for university teaching quality and reformation engineering (No. 2013-2). |
| 文献类型 | 期刊论文 |
| 条目标识符 | http://ir.kiz.ac.cn/handle/152453/9335 |
| 专题 | 科研部门_兽类生态与进化（蒋学龙） 遗传资源与进化国家重点实验室 |
| 通讯作者 | wangyy@mail.sysu.edu.cn |
| 作者单位 | 1.State Key Laboratory of Biocontrol/The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, Guangdong, China 2.Present address: Kadoorie Conservation China, Kadoorie Farm and Botanic Garden, Lam Kam Road, Tai Po, Hong Kong, China 3.State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, Yunnan, China |
| 推荐引用方式 GB/T 7714 | Yang JH,Rao DQ,Wang YY*,et al. A new species of the genus Liuixalus (Anura: Rhacophoridae) from southern China[J]. ZOOTAXA,2015,3990(2):247-258. |
| APA | Yang JH,Rao DQ,Wang YY*,&wangyy@mail.sysu.edu.cn.(2015).A new species of the genus Liuixalus (Anura: Rhacophoridae) from southern China.ZOOTAXA,3990(2),247-258. |
| MLA | Yang JH,et al."A new species of the genus Liuixalus (Anura: Rhacophoridae) from southern China".ZOOTAXA 3990.2(2015):247-258. |
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